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Perona

Japanese name: ペローナ
Nicknamed the Ghost Princess, Perona is a girl dressed in the style of Gothic Lolita and a commander in the army of Gecko Moria. She is also the commander of Perona Wonder Garden, where an army of animal zombies live and protect her. She loots the Thousand Sunny with some zombies and later takes on guard duty while Absalom is attempting to marry Nami. She cannot stand anything that she feels is not cute, such as insects. Perona's dream is to have her own country made up of cute zombie animals with her as their leader. Like many other characters in One Piece, Perona has a unique laugh, which is horohorohoro.

Perona has eaten the Horo Horo no Mi which allows her to create several ghostly apparitions. The ghosts can pass through someone to cause them to go into a sort of induced depression. Other versions of these ghosts can be used as extremely powerful bombs and vary in size. Aside from being able to create ghosts, Perona herself is also able to use ghostly powers such as flight, intangibility, and size manipulation, however, in order to do so Perona must leave her physical body. She is defeated by Usopp, who is the only person immune to her "hollow" attacks, because he claims he is already negative.

She later awakens and decides to flee Thriller Bark in the wake of Oz's rampage. After loading the Straw Hat's ship with food and treasure, she is confronted by Bartholomew Kuma. Perona attempts to capture the Shichibukai for Gecko Moria, but is transported to an unknown location by Kuma.

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